Martin H. Villet

Professor in Entomology

Head: Southern African Forensic Entomology Research Laboratory

PhD (Wits), PGDHE (Rhodes)


Phone +27 [0]46 603 8527

Martin Villet CV

Past Postgraduate Student Supervision

  1. Christian Deschodt (MSc, 2019) - Using four different methods to reach a taxonomic conclusion in dung beetles (Scarabaeinae).
  2. Erica IDS Mbata (MSc, 2018) - The evaluation of potential dietary media, measurement parameters and storage techniques for use in forensic entomotoxicology
  3. Mary Cole (PhD, 2017) - Systematics and biogeography of forest snails, Chondrocyclus (Mollusca: Gastropoda: Caenogastropoda: Cyclophoridae) in southern Africa
  4. Colleen Hepburn (PhD, 2017) - The phenologies of macadamia (Proteaceae) and thrips (Insecta: Thysanoptera) communities in Mpumalanga Province, South Africa
  5. Megan Amy Riddin (PhD, 2017) - The epidemiology of African horse sickness in the Eastern Cape, South Africa
  6. Zanthé Kotzé (MSc, 2016) - Implications and causes of heat accumulation in maggot masses of the sheep blow fly, Lucilia cuprina (Wiedemann) (Diptera: Calliphoridae). (Studied through the University of Pretoria)
  7. Chantal Taylor (MSc, 2016) - Phylogeographical studies of aquatic macroinvertebrate fauna of the southern Great Escarpment. (With distinction)
  8. Lenin Chari (PhD, 2016) - The role of invertebrate aerial predators in cross-boundaries trophic subsidies in the Kowie River catchment.
  9. Sydney Moyo (PhD, 2016) - The role of mayflies and caddisflies in cross-boundaries trophic subsidies in the Kowie River catchment.
  10. Werner Strümpher (PhD, 2016) - Phylogeographic studies of Trogidae (Coleoptera: Scarabaeoidea)
  11. Kirstin Williams (PhD, 2015) – Systematics of Lucilia (Diptera: Calliphoridae) in Africa.
  12. Jaryd Ridgeway (MSc, 2015) - Real-time PCR as a tool in insect pathology.
  13. Isabel Collett (MSc, 2015) - Barcoding of forensically significant South African Beetles.
  14. Megan Riddin (MSc, 2014) - Phylogenetics and biogeography of Mesobola (Pisces: Cyprinidae).
  15. Goetz Neef (MSc, 2014) - Phylogeography of South African forest swallotails (Lepidoptera: Papilionidae).
  16. John M. Midgley (PhD, 2013) – Epigaeic invertebrate biodiversity of the eastern Great Escarpment.
  17. Ashley Kirk-Spriggs (PhD, 2012) - Systematics and biogeography of the Afrotropical Curtonotidae (Diptera).
  18. Helen M. Barber-James (PhD, 2011) - Systematics, morphology, phylogeny and historical biogeography of the mayfly family Prosopistomatidae (Ephemeroptera: Insecta) of the World (Employed at the Albany Museum).
  19. Terence A. Bellingan (MSc, 2011) - Aquatic invertebrate biodiversity in the Tsitsikamma rivers.
  20. Benjamin W. Price (PhD, 2010) - Molecular phylogeny of the southern African platypleurine cicadas.
  21. Nolwazi Mkize (PhD, 2009) - Pests of cultivated (Olea europaea L.) and wild (Olea europea africana) olive trees in the Eastern Cape (Employed at the National Department of Agriculture).
  22. Cameron S. Richards (PhD, 2008) - Thermoecology of blowflies (Diptera: Calliphoridae) in South Africa. (Employed at Natural History Museum, London)
  23. Simon Kamande Kuria (PhD, 2007) - Influence of gall-dwelling ants on invertebrate biodiversity on the swollen-thorn acacia, Acacia drepanolobium in Kenya. (Employed by Walter Sisulu University).
  24. Abraham S. Chawanji (PhD, 2007) - Comparative spermatology and reproductive strategies in cicadas: taxonomic and phylogenetic considerations (Emigrated to New Zealand).
  25. Charles M. Warui (PhD, 2005) - Biomonitoring of grazing by cattle and indigenous herbivores in Kenyan savanna using spiders (Employed by National Museums of Kenya).
  26. Mark Robertson (PhD, 2003) - Preditive modelling of species' potential geographical distributions (Employed by University of Pretoria).
  27. Tamara J. Smith (PhD, 2003) - The Diamondback Moth, Plutella xylostella (L.) (Lepidoptera: Plutellidae) and its biological control in the Eastern Cape province, South Africa (Employed by South African National Biodiversity Institute).
  28. Colleen Hepburn (MSc cum laude, 2008) - Composition and phenology of insect pests in Capsicum (Solanaceae) cultivated in the Makana District, Eastern Cape Province, South Africa. (Independent agricultural consultant)
  29. Kierryn Gendall (MSc, 2008) - Agathis bishopi (Braconidae): its biology and usefulness as a biological control agent for false codling moth (FCM), Cryptophlebia leucotreta, on citrus.
  30. John M. Midgley (MSc, 2008) - Aspects of the thermal ecology of six species of carcass beetles in South Africa. (Continued to PhD)
  31. Unathi L.P. Heshula (MSc, 2005) - Biological control value of Falconia intermedia on Lantana camara in the Eastern Cape (Continued to PhD).
  32. Alfred Stemele (MSc, 2005) - Effects of Bt and intercropping with Artemesia on Diamond-backed Moth (Plutella xylostella) and its parasitoids in small-scale cabbage farms (Registered at UFH).
  33. Sicelo Nofemela (MSc, 2004) - The ecology of certain parasitoids of Plutella xylostella (Employed by Agricultural Research Council).
  34. Nolwazi Mkize (MSc, 2004) - Control of diamondack moth on cabbages by small-scale farmers in the Eastern Cape (Employed by Agricultural Research Council; Continued to PhD).
  35. Nomahlubi Sishuba (MSc, 2004) - Biological control of False Codling Moth (Cryptophlebia leucotreta) in citrus (Employed by Agricultural Research Council; Continued to PhD).
  36. Kirstin A. Williams (MSc, 2003) - Spatial and temporal occurrence of forensically important South African blowflies (Diptera: Calliphoridae) (Employed by Durban Natural Scieince Museum).
  37. Angela Bownes (MSc, 2003) - The structure of ant communities and their impact on soil-pupating pests in citrus orchards in the Grahamstown area of the Eastern Cape (Employed by Agricultural Research Council; continued to PhD).
  38. Nicola Lunt (MSc, 2003) - Applied studies of some Southerm African blowflies (Diptera: Calliphoridae) of forensic importance (Employed at Marwell Trust; continued to PhD).
  39. Satch Moshiane (MSc, 2002) - The ecology of Diamond-backed Moth (Plutella xylostella) in canola crops (Employed by Agricultural Research Council).
  40. Jennifer Youthed (MSc, 1997) - A predictive biogeography for selected alien plant invaders in South Africa. (Employed by Eastern Cape Government).

 Selected Forensic Research Publications

  1. Campobasso CP, Bugelli V, Carfora A, Borriello R, Villet MH (in prep) Advances in entomotoxicology: weaknesses and strengths. In Byrd J, Tomberlin J. (Eds.) Forensic Entomology: The Utility of Arthropods in Legal Investigations (3rd Ed).
  2. Badenhorst R, Villet MH. 2018. The uses of Chrysomya megacephala (Fabricius, 1794) (Diptera: Calliphoridae) in forensic entomology. Forensic Sciences Research 3: 2-15. doi: 10.1080/20961790.2018.1426136
  3. Villet MH. 2017. Forensic significance of Diptera. In: Kirk-Spriggs, A.H. & Sinclair, B.J., eds, Manual of Afrotropical Diptera. Volume 1. Introductory chapters and keys to Diptera families. Suricata 4. Pretoria: SANBI Graphics & Editing, 175–193 pp.
  4. Daniel CA, Midgley JM, Villet MH. 2017. Identification of larvae of the Afrotropical species of Thanatophilus Leach, 1817 (Coleoptera, Silphidae). African Invertebrates 58: 1-10. DOI: 10.3897/afrinvertebr.58.12966 Corrigenda:
  5. da Silva E, Wilhelmi B, Villet MH. 2017. Forensic entomotoxicology revisited - towards professional standardization of study designs. International Journal of Legal Medicine. 131: 1399-1412. DOI: 10.1007/s00414-017-1603-9
  6. Lutz L, Williams KA, Villet MH, Ekanem M, Szpila K. 2017. Species identification of African blowflies (Diptera: Calliphoridae) of forensic importance. International Journal of Legal Medicine 132: 831–842. DOI: 10.1007/s00414-017-1654-y
  7. Villet MH, Clitheroe C-L, Williams KA. 2017. Forensic implications of the temporal occurrence of flesh flies (Diptera: Sarcophagidae) at carrion-baited traps in Grahamstown, South Africa. African Invertebrates 58: 1-8. DOI: 10.3897/AfrInvertebr.58.9537.
  8. Williams KA, Wallman JF, Lessard BD, Kavazos CRJ, Mazungula DN, Villet MH. 2017. Nocturnal oviposition behavior of carrion flies (Diptera: Calliphoridae, Sarcophagidae) in the southern hemisphere (South Africa and Australia) and its forensic implications. Forensic Science, Medicine, and Pathology. 13: 123-134. DOI: 10.1007/s12024-017-9861-x
  9. Kotzé Z, Villet MH, Wheldon CW. 2016. Heat accumulation and development rate of massed maggots of the sheep blowfly, Lucilia cuprina (Diptera: Calliphoridae). Journal of Insect Physiology 95: 98-104. DOI: 10.1016/j.jinsphys.2016.09.009
  10. STRÜMPHER WP, SOLE CL, VILLET MH & SCHOLTZ CH. 2016. Allopatric speciation in the flightless Phoberus capensis (Coleoptera: Trogidae) group, with description of two new species. Insect Systematics & Evolution DOI: 10.1163/1876312X-47022138.
  11. STRÜMPHER WP, VILLET MH, SOLE CL, SCHOLTZ CH. 2016. Overview and revision of the extant genera and subgenera of Trogidae (Coleoptera: Scarabaeoidea). Insect Systematics & Evolution 47: 53 – 82. DOI: 10.1163/1876312X-46052133.
  12. WILLIAMS KA, VILLET MH, LAMB J. 2016. Phylogenetic radiation of the greenbottle flies (Diptera, Calliphoridae, Luciliinae). ZooKeys 568: 59-86. DOI: 10.3897/zookeys.568.6696.
  13. Strümpher WP, Villet MH, Sole CL, Scholtz CH. 2016. Overview and revision of the extant genera and subgenera of Trogidae (Coleoptera: Scarabaeoidea). Insect Systematics & Evolution 47: 53 – 82. DOI: 10.1163/1876312X-46052133.
  14. Williams KA, Villet MH, Lamb J. 2016. Phylogenetic radiation of the greenbottle flies (Diptera, Calliphoridae, Luciliinae). ZooKeys 568: 59-86. DOI: 10.3897/zookeys.568.6696
  15. Kotzé Z, Villet MH, Weldon CW. 2015. Effect of temperature on development of the blowfly, Lucilia cuprina (Wiedemann) (Diptera: Calliphoridae). International Journal of Legal Medicine 129: 1155-1162. DOI 10.1007/s00414-015-1153-y
  16. AMENDT J, CAMPOBASSO CP, DADOUR IR, GAUDRY E, HALL MJR, VILLET MH, MORETTI TC, ANDERSON GS, SUKONTASON KL, 2015. Standard practices. In: Tomberlin JK, Benbow ME, eds. International Dimensions and Frontiers in Forensic Entomology. Boca Raton: Taylor & Francis. DOI: 10.1201/b18156-35
  17. VILLET MH, 2015. History, accomplishments, and challenges of forensic entomology in Africa. In: Tomberlin JK, Benbow ME, eds. International Dimensions and Frontiers in Forensic Entomology. Boca Raton: Taylor & Francis. DOI: 10.1201/b18156-20
  18. WILLIAMS KA, VILLET MH, 2014. Morphological identification of Lucilia sericata, Lucilia cuprina and their hybrids (Diptera, Calliphoridae). ZooKeys 420:69-85.
  19. WILLIAMS KA, RICHARDS CS, VILLET MH, 2014. Predicting the geographic distribution of Lucilia sericata and Lucilia cuprina (Diptera: Calliphoridae) in southern Africa. African Invertebrates 55: 157-170.
  20. STRÜMPHER WP, SOLE CL, VILLET MH, SCHOLTZ CH, 2014. Phylogeny of the family Trogidae (Coleoptera: Scarabaeoidea) inferred from mitochondrial and nuclear ribosomal DNA sequence data. Systematic Entomology 39: 548-562.
  21. RIDGEWAY JA, MIDGLEY JM, COLLETT IJ, VILLET MH, 2014. Advantages of using development models of the carrion beetles Thanatophilus micans (Fabricius) and T. mutilatus (Castelneau) (Coleoptera: Silphidae) for estimating minimum post mortem intervals, verified with case data. Int. J. Legal Med. 128: 207-220.
  22. WILLIAMS KA, VILLET MH, 2013. Ancient and modern hybridization between Lucilia sericata and Lucilia cuprina (Diptera: Calliphoridae). European Journal of Entomology 110: 187-196.
  23. MARINHO MAT, JUNQUEIRA ACM, PAULO DF, ESPOSITO MC, VILLET MH, AZEREDO-ESPIN AML, 2012. Molecular phylogenetics of Oestroidea (Diptera: Calyptratae) with emphasis on Calliphoridae: insights into the inter-familial relationships and additional evidence for paraphyly among blowflies. Mol. Phylog. Evol. 65: 840-854.
  24. PICARD CJ, VILLET MH, WELLS JD, 2012. Amplified fragment length polymorphism confirms reciprocal monophyly in Chrysomya putoria and Chrysomya chloropyga: a correction of reported shared mtDNA haplotypes. Med. Vet. Entomol. 26: 116-119.
  25. VILLET MH, 2011. African carrion ecosystems and their insect communities in relation to forensic entomology: a review. Pest Technology 5: 1-15.
  26. VILLET MH, AMENDT J, 2011. Advances in entomological methods for estimating time of death. In: Turk EE, ed. Forensic Pathology Reviews. Heidelberg: Humana Press. in press.
  27. SZPILA K, VILLET MH, 2011. Morphology and identification of first instar larvae of African blowflies (Diptera: Calliphoridae) commonly of forensic importance. Med. Vet. Entomol.48:738-752.
  28. MIDGLEY JM, RICHARDS CS, VILLET MH, 2010. The utility of Coleoptera in forensic investigations. In: Amendt J, Campobasso CP, Goff ML, Grassberger M, eds. Current Concepts in Forensic Entomology. Heidelberg: Springer, pp. 57-68.
  29. VILLET MH, RICHARDS CS, MIDGLEY JM, 2010. Contemporary precision, bias and accuracy of minimum post-mortem intervals estimated using development of carrion-feeding insects. In: Amendt J, Campobasso CP, Goff ML, Grassberger M, eds. Current Concepts in Forensic Entomology. Heidelberg: Springer, pp. 109-137.
  30. MIDGLEY JM, VILLET MH, 2009. Development of Thanatophilus micans (Fabricius 1794) (Coleoptera: Silphidae) at constant temperatures. Int. J. Legal Med.123:285-292.
  31. MIDGLEY JM, VILLET MH, 2009. Effect of the killing method on post-mortem change in length of larvae of Thanatophilus micans (Fabricius, 1794) (Coleoptera: Silphidae) stored in 70% ethanol. Int. J. Legal Med.123:103-108.
  32. RICHARDS CS, CROUS KL, VILLET MH, 2009. Models of development for the blow fly sister species Chrysomya chloropyga and C. putoria. Med. Vet. Entomol.23:56-61.
  33. RICHARDS CS, VILLET MH, 2009. Data quality in thermal summation models of development of forensically important blowflies (Diptera: Calliphoridae): a case study. Med. Vet. Entomol.23:269-276.
  34. RICHARDS CS, PRICE BW, VILLET MH, 2009. Thermal ecophysiology of seven carrion-feeding blowflies in Southern Africa. Ent. Exp. Appl.131:11-19.
  35. RICHARDS CS, WILLIAMS KA, VILLET MH, 2009. Predicting geographic distribution of seven blowfly species (Diptera: Calliphoridae) in South Africa. Afr. Entomol.17:170-182.
  36. TOURLE RA, DOWNIE DA, VILLET MH, 2009. Flies in the ointment: a morphological and molecular comparison of Lucilia cuprina and L. sericata (Diptera: Calliphoridae) in South Africa. Med. Vet. Entomol.23:6-14.
  37. HARVEY ML, GAUDIERI S, VILLET MH, DADOUR IR, 2008. A global study of forensically significant calliphorids: implications for identification. Forensic Sci. Int.177:66-76.
  38. RICHARDS CS, PATERSON ID, VILLET MH, 2008. Estimating the age of immature Chrysomya albiceps (Diptera: Calliphoridae), correcting for temperature and geographical latitude. Int. J. Legal Med.122:271-279.
  39. RICHARDS CS, VILLET MH, 2008. Factors affecting accuracy and precision of thermal summation models of insect development used to estimate postmortem intervals. Int. J. Legal Med.122:401-408.
  40. VILLET MH, 2007. An inexpensive geometrical micrometer for measuring small, live insects quickly without harming them. Ent. Exp. Appl.122:279-280.
  41. WILLIAMS KA, VILLET MH 2006. A history of southern African research relevant to forensic entomology. South African Journal of Science 102, 59-65.
  42. WILLIAMS KA, VILLET MH, 2006. A new and earlier record of Chrysomya megacephala in South Africa, with notes on another exotic species, Calliphora vicina (Diptera: Calliphoridae). Afr. Invert. 47:347-350.
  43. VILLET MH, MACKENZIE B, MULLER WJ, 2006. Larval development of the carrion-breeding flesh fly Sarcophaga (Liosarcophaga) tibialis Macquart (Diptera: Sarcophagidae) at constant temperatures. Afr. Entomol. 14:357-366.
  44. HARVEY ML, MANSELL MW, VILLET MH, DADOUR IR, 2003. Molecular identification of some forensically important blowflies of southern Africa and Australia. Med. Vet. Entomol. 17:363-369.
  45. MUSVASVA E, WILLIAMS KA, MULLER WJ, VILLET MH, 2001. Preliminary observations on the effects of hydrocortisone and sodium methohexital on development of Sarcophaga (Curranea) tibialis Macquart (Diptera: Sarcophagidae), and implications for estimating post mortem interval. Forensic Sci. Int. 120:37-41.

Last Modified: Fri, 17 Apr 2020 11:06:27 SAST