Prof Alan Hodgson

Alan Hodgson 120x200

PhD Manchester University (1980)
DSc Manchester University (2004)

Phone: + 27 (0) 46 603 7093
Email: a.hodgson 'at'


  • Invertebrate gametogenesis, gamete structure and function
  • Invertebrate reproduction
  • Molluscan biology
  • Biology of limpets
  • Biology and ecology of estuarine invertebrates
  • Knysna Basin Project

Editorial Work

Co-editor, Invertebrate Reproduction & Development

Associate Editor, African Journal of Aquatic Science


Current students

James Radloff (M.Sc.) - Recruitment of the invasive bivalve Mytilus galloprovincicalis in the Knysna Estuary.

Murray Roodt (M.Sc.) – Invertebrate biodiversity on floating docks in Thesen Islands Marina, Knysna.

Nina de Villers (M.Sc.) - Compaison of the fauna of natural and artifical substrata in the Keurbooms Estuary. 

Cassandra Barker (Honours) - Population structure of the Knysna seahorse in Codium tenue.

David Taylor (Honours) - Comparison of fish diversity using baited and unbaited action cameras in two permanently open estuaries.

Recently Graduated students:

Melissa Pollard (M.Sc.) – Utilization of sea grass meadows by fish in the Knysna estuary.

Louw Claassens (Ph.D.) - Biology and ecology of the Knysna seahorse.



Some recent publications

HODGSON AN, DICKENS J. 2012. Activity of the mangrove snail Cerithidea decollata (Gastropoda: Potamididae) in a warm temperate South African estuary. Estuarine and Coastal Shelf Science. 109: 98-106.

HODGSON AN, FERENZ H-J, SCHNEIDER S. 2013. Formation of sperm bundles in Pterostichus nigrita (Coleoptera: Carabidae). Invertebrate Reproduction and Development.57: 120-131.

HODGSON AN, ECKELBARGER KJ, HODGSON V, YOUNG CM. 2013. Spermatozoon structure of Acesta oophaga (Limidae), a cold-seep bivalve. Invertebrate Reproduction and Development. 57: 70-73.

KRUGER S, FERENZ H-J, RANDALL M, HODGSON AN. 2014. Structure of the male reproductive accessory glands of Pterostichus nigrita (Coleoptera: Carabidae), their role in spermatophore formation. Invertebrate Reproduction and Development. 58: 75-88.

ECKELBARGER KJ, HODGSON AN. 2014. Ultrastructure of spermiogenesis and synthesis of enigmatic cytoplasmic inclusions in the spirally-shaped spermatozoon of the polychaete Spio setosa (Annelida: Spionidae). Invertebrate Biology. 133: 146-157.

HODGSON AN, BOOTH AJ, DAVID-ENGELBRECHT V, HENNINGER TO. 2014. Some life-history parameters of the non-native amphipod Platorchestia platensis (Talitridae) in a warm temperate South African estuary. Transactions of the Royal Society of South Africa. 69: 97-106.

NYALUNGU NP, HODGSON AN, FRONEMAN PW, DOPOLO M, MASUBELELE M. 2014. Bio- and physic-chemical factors affecting the temporal and spatial variability in the distribution of Palaemon peringueyi (Decapoda: Crustacea) in the permanently open Kariega Estuary, South Africa. African Journal of Aquatic Science 39: 435-442.

PINCHUCK SC, ALLANSON, BR AND HODGSON AN. 2015. Evolutionary retention of defensive lateral pedal glands in the smallest siphonariid limpet (Gastropoda: Pulmonata). African Zoology. 50 (4): 327-330.

POLLARD M, HODGSON AN. 2016. Mytilus galloprovincialis (Mollusca: Bivalvia) in a warm-temperate South Africa estuarine embayment. African Journal of Aquatic Science 41: 57-65.

HODGSON AN, RIDGEWAY JA, VILLET MH. 2016. Sperm ultrastructure and spermatodesm morphology of the spittle bug Locris transversa (Thunberg 1822) (Hemiptera: Cercopidae). Invertebrate Reproduction and Development 60: 87-94.

DIEMAR N, GRIFFITHS CL, HODGSON AN. 2016. Disentangling the identities and distribution patterns of the introduced beachfleas Orchestia gammarellus and Platorchestia platensis (Crustacea: Amphipoda: Talitridae) in South Africa.  African Zoology 51: 203-210.

POLLARD M, HODGSON AN, KOK HM, WHITFIELD AK. 2017. Eelgrass beds and bare substrata – sparid and mugilid composition in contrasting littoral estuarine habitats. African Journal of Marine Science 39: 211-224.


CLAASSENS L, BOOTH AJ, HODGSON AN. 2018. An endangered seahorse selectively chooses and artificial structure. Environmental Biology of Fishes 101: 723-733.

CLAASSENS L, HODGSON AN. 2018. Gaining insights into in situ behaviour of an endangered seahorse using action cameras. Journal of Zoology 304: 98-108.

CLAASSENS L, HODGSON AN. 2018. Monthly population density and structure patterns of an endangered seahorse Hippocampus capensis: a comparison between natural and artificial habitats. Journal of Fish Biology 92: 2000-2015.

POLLARD M, WHITFIELD AK, HODGSON, AN. 2018. Possible influences of a macroalgal bloom in eelgrass beds on fish assemblages in the lower Knysna Estuary, South Africa. African Journal of Aquatic Science 43: 319-323.

PINCHUCK SC, HODGSON AN. 2018. Retinal ultrastructure of patellogastropods revisited. Journal of Molluscan Studies 84: 494-497.

Book Chapter

HODGSON, AN. 2018. Spermatogenesis and spermiogenesis in the Mollusca. In M.K. Skinner (ed.), Encyclopaedia of Reproduction. Vol. 6, pp336-343. Academic Press.


Last Modified: Thu, 01 Nov 2018 15:52:51 SAST